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For those who are interested I suggest getting a copy of the following article in Alytes

Andreone, Franco; Giacoma, Cristina; Guarino, Fabio M.; Mercurio, Vincenzo; Tessa, Giulia; 2011; Age profile in none Mantella poison frogs from Madagascar, as revealed by skeletochronological analysis; Alytes, 27 (3):73-84

It is interesting as it gives sexual maturity in wild frogs and life span in wild frogs. The species studied were M. baroni, M. bernhardi, M. sp. aff. expectata; M. cowani, M. crocea, M. laevigata, M. nigricans, M. pulchra and M. viridis.

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